CO grants status 24 hours after the flight segment where you met the mileage requirement. When I first qualified for Silver on the first leg, I did not get the Elite mileage bonus on the second leg, but I did on a flight a few days later.
A flight taken Tuesday
should post by Friday if it's on CO metal. However, sometimes they are slow to post...go figure!
If the flight hasn't posted by Friday, show up at the airport, explain that the critical miles haven't posted yet, show them your last mileage statement and your boarding pass from Tuesday's flight...and beg...very politely! If the counter agent says no, then ask the President's Club agent and/or the gate agent. That worked for me, but there are no guarantees. All of this assumes, of course, that there are upgrades available.
Two hours prior to flight time, seat assignments are transferred from reservations to airport control. And the airport agents have far more discretion with seat assignments than reservation agents.
Again, if you don't like the answer the first agent gives you, ask someone else!
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In Economy...no one can hear you scream!
http://www.denycontinentalthefreddie.com/